Jason Lee

Dave

Justin Long

Alvin (voice)

Matthew Gray Gubler

Simon (voice)

Jesse McCartney

Theodore (voice)

Kimberly Williams-Paisley

Samantha

Josh Green

Miles

Tony Hale

Agent Suggs

Bella Thorne

Ashley Grey

Eddie Steeples

Barry

Christina Applegate

Brittany (voice)

Kaley Cuoco

Eleanor (voice)

Anna Faris

Jeanette (voice)